One of Thriller Bark's Mysterious Four. Endowed with the powers of the Hollow-Hollow Fruit, she commands the Wild Zombies and the Surprise Zombies. Serves under Gecko Moria.

Type Class 1 Class 2 Rarity Cost
DEX Shooter Driven 3 10
Power Sockets Combo Price Max Lv.(Exp.)
1 4 300 50(392,109)
Lv. HP Attack RCV
Base 1 134 44 55
Max 50 944 422 222
Special Negative Hollow
Description Reduces damage taken by 90%, drastically reduces enemies’ defense, and deals spirit poison damage for 1 turn
Captain Ability Perona’s Mysterious Ghost
Description Boosts DEX characters’ RCV by 1.5x
Pre-Evolution Character Evolvers Needed Post-Evolution
Perona Sea Horse
Green Pirate Penguin
Green Hermit Crab
Green Armored Crab
Green Striped Dragon
Ghost Princess Perona

Where to find Evolvers

Tandem Attacks

Tandem Attacks Description
Character 1 None
Character 2 None
Character 3 None
Character 4 None
Character 5 None

Characters by Type:
Strategy Guide Quest Guide Character List Characters by Type Tandem Attacks Evolutions at a Glance Finding Evolvers Ships Official Site